Refereed publications


Titlyanov EA, Titlyanova TV & Tokeshi M (2015). Coral-algal relations in reef systems: on the positive roles of algae in reef recovery processes. Coastal Ecosystems, 2, 1-20.

Susanto HA, Suraji & Tokeshi M (2015). Management of coral reef ecosystems in Indonesia: past, present and the future. Coastal Ecosystems, 2, 21-41.

Arakaki S, Kawabata Y & Tokeshi M(2015). Assessing the feasibility of bio-logging research in adult temperate bass, Lateolabrax latus, in subtropical-temperate coastal waters of southwestern Japan. Coastal Ecosystems, 2, 42-52.

Arakaki S, Tsuchiya M & Tokeshi M (2014). Testing latitudinal patterns of tidepool fish assemblages: local substrate characteristics affect regional-scale trends. Hydrobiologia (Springer) 733: 45-62.

Yoko-o M & Tokeshi M (2014). Pronounced difference in community structure between the edge and the interior: analysing small-scale variability of a maritime woodland. Coastal Ecosystems, 1, 54-65.

Arakaki S, Hutchinson N & Tokeshi M(2014). Foraging ecology of a large opportunistic predator (adult Lateolabrax latus) on a temperate-subtropical rocky shore. Coastal Ecosystems, 1, 14-25.

Tokeshi M, Arakaki S & Daud JRP (2013). Consuming diversity: analysis of seasonal catch patterns in multi-species artisanal reef fisheries in North Sulawesi, eastern Indonesia.DPacific Science, 67, 1-13.

Arakaki S & Tokeshi M (2012). Species and size matter: an experimental study of microhabitat use under the influence of competitive interactions in intertidal gobiids. Journal of Experimental Marine Biology and Ecology, 418-419, 59-68.

Yoko-o M. & Tokeshi M. (2012). Assessing context-dependent survival of Quercus glauca seeds in a humid maritime woodland.  Plant Ecology, 213: 143-153. pdf(1032k)

Tokeshi, M & Arakaki, S. (2012). Habitat complexity in aquatic systems: fractals and beyond.  Hydrobiologia, 685: 27-47. pdf(1376k) "invited review -IMPORTANT"

Tokeshi, M & Arakaki, S. (2011). Continuous in-situ observation of reef-associated EchinodermataDGalaxea, 13, 21-22.

Tokeshi, M. (2011). Spatial structures of hydrothermal vents and vent-associated megafauna in the back-arc basin system of the Okinawa Trough, western Pacific.  Journal of Oceanography, 67, 651-665. pdf(3235k) (Important photos)

Tokeshi, M. & Daud, J.R.P. (2011). Feeding electivity in Acanthaster planci: a null model analysis.  Coral Reefs, 30, 227-235. pdf(707k)

Yoko-o M. & Tokeshi M. (2011). Morphological variation along the sea-land gradient: trees in a subtropical maritime woodland. Journal of Forest Research, 16, 55-61. pdf(488k)

Arakaki, S. & Tokeshi, M (2011). Analysis of spatial niche structure in coexisting tidepool fishes: null models based on multi-scale experiments.  Journal of Animal Ecology, 80, 137-147. pdf(1032k)

Tokeshi, M. & Tanaka, K. (2010). Dominance of tabular Acroporid species and the abundance of echinoid grazers in high-latitude coral assemblages of Amakusa, southwestern Japan.  Galaxea, 12, 87. pdf(1597k)

Kurimoto, M. & Tokeshi M. (2010). Variation on a theme of herbivory: Corallina- hermit crab relationship on a temperate-subtropical rocky shore. Oikos, 119, 1401-1408. pdf(467k)

Nozawa Y, Tokeshi M, Nojima S (2008). Structure and dynamics of a high-latitude scleractinian coral community in Amakusa, southwestern Japan. Marine Ecology Progress Series 358: 151-160

Tokeshi, M. & Arakaki, S. (2007).  On estimating diversity in species-rich fish assemblages: an example with coral reef fishes of eastern Indonesia. Journal of Fish Biology, 71, 1879-1887.

Kawai,T & Tokeshi, M. (2007).  Testing the facilitation-competition paradigm under the stress-gradient hypothesis: decoupling multiple stress factors.  Proceedings of the Royal Society, London, B, 274, 2503-2508. 

Londono-Cruz, E. & Tokeshi, M. (2007).  Testing scale-variance in species-area and abundance-area relationships in a local assemblage: an example from a subtropical boulder shore.  Population Ecology, 49, 275-285. pdf(625k)

Tokeshi, M
., Yoko-o, M., Daud, J.R.P. & Domits, M. (2007).  Hypolycaena erylus feeding on the mangrove apple and attended by Oecophylla weaver ants, in North Sulawesi, Indonesia (Lepidoptera: Lycaenidae).  Tropical Lepidoptera, 17, 34-35. pdf(1293k)

Tokeshi, M
& Yoko-o, M. (2007).  New record of the tropical swallowtail moth Lyssa zampa (Butler) (Lepidoptera: Uraniidae) from mainland Japan.  Entomological Science, 10, 103-106.

Arakaki, S & Tokeshi, M (2006).  Short-term dynamics of tidepool fish community: diel and seasonal variation.  Environmental Biology of Fishes, 76, 221-235.

Kawai,T & Tokeshi, M. (2006).  Asymmmetric coexistence: bidirectional positive and negative interactions between goose-barnacles and mussels.  Journal of Animal Ecology, 75, 928-941.

Nozawa, Y., Tokeshi, M & Nojima, S (2006)  Reproduction and recruitment of scleractinian corals in a high-latitude coral community, Amakusa, southwestern Japan.  Marine Biology, 149, 1047-1058.

Arakaki, S. & Tokeshi, M. (2005). Microhabitat selection in intertidal gobiid fishes: species and size-associated interaction. Marine Biology Research 1:39-47

Taniguchi, H. & Tokeshi, M. (2004). Effects of habitat complexity on benthic assemblages in a variable environment. Freshwater Biology, 49, 1164-1178.pdf(364k)

Kawai, T. & Tokeshi, M. (2004). Variable modes of facilitation in the upper intertidal: goose barnacles and mussels. Marine Ecology Progress Series, 272, 203-213.

Paruntu, C.P. & Tokeshi, M. (2003). Variability in the reproductive characteristics of local populatiopns of an intertidal gastropod, Nerita japonica (Dunker). Benthos Research, 58, 7-14.

Taniguchi, H., Nakano, S & Tokeshi, M. (2003). Influences of habitat complexity on the diversity and abundance of epiphytic invertebrates on plants. Freshwater Biology, 48, 718-728. pdf(646k)

Tokeshi, M. & Schmid, P.E. (2002) Niche division and abundance: an evolutionary perspective. Population Ecology, 44, 189-200.

Schmid, P.E., Tokeshi, M. & Schmid-Araya, J.M. (2002). Scaling in stream communities. Proceedings of the Royal Society, London, Series B, 269, 2587-2594.

Tokeshi, M. (2002) Spatial distribution of a deep-sea crinoid Pentametrocrinus tuberculatus in the Izu-Ogasawara Arc, western Pacific.  Journal of Zoology, (London), 258, 291-298.

Ota, N. & Tokeshi, M. (2002) A population study of two carnivorous buccinids (Gastropod:Buccinidae) on an intertidal stony shore. Venus, 60, 261-271.

Romero, L. & Tokeshi, M. (2002) Variabilidad fenotipica de Nodilittorina en la Reserva de Paracas (Phenotypic variability of Nodilittorina in the Paracas Reserve). Revista Peruana de Biologia, 9, 39-48.

Thamrin, Nojima, S. & Tokeshi, l. (2001). An experimental analysis of planula release in a scleractinian coral Alveopora japonica. Galaxea, 3, 25-30.

Thamrin, Tokeshi, M. & Nojima, S. (2001). Effect of coral-inhabiting barnacle (Cantellius pallidus) on planula production in a scleractinian coral Alveopora japonica.  Ophelia, 55, 93-100.


Tioho,H., Tokeshi, M.
& Nojima, S. (2001).  Experimental analysis of recruitment in a scleractinian coral at high latitude.  Marine Ecology Progress Series, 213, 79-86.


Tokeshi, M. (2001)  Coexistence.  In: Encyclopedia of Life Sciences, online edition, Macmillan Publishers, London.

Tokeshi, M. & Romero, L. (2000). Spatial overlap and coexistence in a mussel-associated polychaete assemblage on a South American rocky shore. PSZN Marine Ecology,  21, 247-261.

Schmid, P.E., Tokeshi, M. & Schmid-Araya, J.M. (2000). Relation between population density and body size in stream communities. Science, 289, 1557-1560. pdf (276k)

Ota, N. & Tokeshi, M. (2000b). Population analysis of Ruditapes variegatus (Sowerby) (Bivalvia: Veneridae) on an intertidal boulder shore. Venus, 59, 29-36.

Ota, N. & Tokeshi, M. (2000a). A comparative study of feeding and growth in two coexisting species of carnivorous gastropods. Marine Biology, 136, 101-114.

Tokeshi, M., Ota, N. & Kawai, T. (2000). A comparative study of morphometry in shell-bearing molluscs. Journal of Zoology (London), 251, 31-38. pdf (136k)

Kadono, A, Takemon, Y & Tokeshi, M. (1999). Mobility and habitat colonization in stream invertebrates: an experimental study. Japanese Journal of Limnology, 60, 215-222.

Arakaki, S., Yamahira, K. & Tokeshi, M. (1999). Sex change and spatial distribution pattern in an intertidal holothurian Polycheira rufescens in the reproductive season. Researches on Population Ecology, 41, 235-242.

Tokeshi, M. (1999). SPECIES COEXISTENCE: Ecological and Evolutionary Perspectives. Blackwell Science, Oxford.

Tokeshi, M. & Reinhardt, K.(1996). Reproductive behaviour in Chironomus anthracinus (Diptera: Chironomidae), with a consideration of the evolution of swarming. Journal of Zoology, (London), 240, 103-112.

Tokeshi, M. (1996b). Power Fraction: a new explanation of relative abundance patterns in species-rich assemblages. Oikos, 75, 543-550.pdf(2806k)

Tokeshi, M. (1996a). Species coexistence and abundance: patterns and processes. Ecological Perspective of Biodiversity (Ed. by T. Abe, S. Levin and M. Higashi), Springer-Verlag, New York.

Tokeshi, M. & Romero, L. (1995b). Quantitative analysis of foraging behaviour in a field population of the South American sun-star, Heliaster helianthus. Marine Biology, 122, 297-303.

Tokeshi, M. & Romero, L. (1995a). Filling a gap: dynamics of space occupancy on a mussel-dominated subtropical rocky shore. Marine Ecology Progress Series, 119, 167-176.

Tokeshi, M. (1995f). On the mathematical basis of the variance-mean power relationship. Researches on Population Ecology, 37, 43-48.

Tokeshi, M. (1995e). Polychaete abundance and dispersion patterns: a non-trivial 'infaunal' assemblage on the Pacific South American rocky shore. Marine Ecology Progress Series, 125, 137-147.

Tokeshi, M. (1995d). Randomness and aggregation: analysis of dispersion in an epiphytic chironomid community. Freshwater Biology, 33, 567-578.

Tokeshi, M. (1995c). Species interactions and community structure. Chironomidae: biology and ecology of non-biting midges (Ed. by Armitage et al.), pp. 297-335. Chapman & Hall, London.

Tokeshi, M. (1995b). Production ecology. Chironomidae: biology and ecology of non-biting midges (Ed. by Armitage et al.), pp. 269-296. Chapman & Hall, London.

Tokeshi, M. (1995a). Life cycles and population dynamics. Chironomidae: biology and ecology of non-biting midges (Ed. by Armitage et al.), pp. 225-268. Chapman & Hall, London.

Takacs, V. & Tokeshi, M. (1994). Spatial distribution of two chironomid species in the bottom sediment of Lough Neagh, Northern Ireland. Aquatic Insects, 16, 125-132.

Tokeshi, M. & Osawa, N. (1994). Niche relations in insect herbivore assemblages. Individuals, Populations and Patterns in Ecology (Ed. by S.R. Leather, A.D.Watt, N.J. Mills and K.F.A. Walters), pp. 309-318. Intercept Press, Andover, England.

Tokeshi, M. (1994b). Biodiversity in aquatic systems: some conceptual and theoretical issues. In: Biodiversity: Its Complexity and Role (Ed. by M. Yasuno and M.M. Watanabe), pp. 211-225. Global Environmental Forum, Tokyo.

Tokeshi, M. (1994a). Community ecology and patchy freshwater habitats. Aquatic Ecology: Scale, Pattern and Process. (Ed. by P.S. Giller, A.G. Hildrew and D.G.Raffaelli), pp. 63-91. Blackwell Scientific Publications, Oxford.

Takacs, V. and Tokeshi, M. (1993). Feeding habits of Procladius signatus in Lough Neagh, Northern Ireland. Irish Naturalists' Journal, 24, 282-283.

Tokeshi, M. (1993c). On the evolution of commensalism in the Chironomidae. FreshwaterBiology, 29, 481-489.

Tokeshi, M. (1993b). Species abundance patterns and community structure. Advances in Ecological Research, 24, 111-186. pdf(5293k)

Tokeshi, M. (1993a). The structure of diversity in an epiphytic chironomid community. Netherlands Journal of Aquatic Ecology, 26, 461-470.

Tokeshi, M. (1992). Dynamics of distribution in animal communities: theory and analysis. Researches on Population Ecology, 31, 279-293. pdf(1367k)

Tokeshi, M. (1991e). Graphical analysis of predator feeding strategy and prey importance. Freshwater Forum, 1, 79-83. pdf(5244k)

Tokeshi, M. (1991d). Faunal assembly in chironomids (Diptera): generic association and spread. Biological Journal of the Linnean Society, 44, 353-367.

Tokeshi, M. (1991c). Extraoral and intraoral feeding: flexible foraging tactics in the South American sun-star, Heliaster helianthus. Journal of Zoology, 225, 339-347.

Tokeshi, M. (1991b). Aggregation, habitat characteristics and species coexistence. Verhandlungen der Internationale Vereinigung fur Theoretische und Angewandte Limnologie, 24, 2964.

Tokeshi, M. (1991a). On the feeding habits of Thienemannimyia festiva (Diptera: Chironomidae). Aquatic Insects, 13, 9-16.

Tokeshi, M. (1990c). Density-body size allometry does not exist in a chironomid community on Myriophyllum. Freshwater Biology, 24, 613-618.

Tokeshi, M. (1990b). Chironomid fauna of Lough Neagh, Northern Ireland (Diptera; Chironomidae). Entomologist's Gazette, 41, 145-148.

Tokeshi, M. (1990a). Niche apportionment or random assortment: species abundance patterns revisited. Journal of Animal Ecology, 59, 1129-1146. pdf(1273k)

Tokeshi, M., Estrella, C. & Paredes, C. (1989). Feeding ecology of a size-structured predator population, the South American sun-star Heliaster helianthus. Marine Biology, 100, 495-505.

Tokeshi, M., Romero, L. & Tarazona, J. (1989). Spatial coexistence of mussel-associated, free-ranging polychaetes in a subtropical intertidal habitat. Journal of Animal Ecology, 58, 681-692.

Tokeshi, M. (1989). Development of a foraging model for a field population of the South American sun-star, Heliaster helianthus. Journal of Animal Ecology, 58, 189-206.

Tokeshi, M., Estrella, C. & Tarazona, J. (1988). Estudio preliminar de las relaciones predador-presas en Heliaster helianthus (Asteroidea: Echinodermata). Recursos y Dinamica del Ecosistema de Afloramiento Peruano (Ed. by H. Salzwedel and A. Landa), pp.141-145, Volumen extraordinario del Boletin del IMARPE.

Tokeshi, M. (1988). Two commensals on a host: habitat partitioning by a ciliated protozoan and a chironomid on the burrowing mayfly Ephemera danica. Freshwater Biology, 20, 31-40.

Tokeshi, M. & Townsend, C.R. (1987). Random patch formation and weak competition: coexistence in an epiphytic chironomid community. Journal of Animal Ecology, 56, 833-845.

Tokeshi, M. & Pinder, L.C.V. (1986). Dispersion of epiphytic chironomid larvae and the probability of random colonization. Internationale Revue der gesamten Hydrobiologie, 71, 613-620.

Tokeshi, M. (1986c). Resource utilization, overlap and temporal community dynamics : a null model analysis of an epiphytic chironomid community. Journal of Animal Ecology, 55, 491-506.

Tokeshi, M. (1986b). Population dynamics, life histories and species richness in an epiphytic chironomid community. Freshwater Biology, 16, 431-442.

Tokeshi, M. (1986a). Population ecology of the commensal chironomid Epoicocladius flavens on its mayfly host Ephemera danica. Freshwater Biology, 16, 235-243.

Tokeshi, M. & Pinder, L.C.V. (1985). Microhabitats of stream invertebrates on two submersed macrophytes with contrasting leaf morphology. Holarctic Ecology, 8, 313-319.

Tokeshi, M. (1985). Life-cycle and production of the burrowing mayfly, Ephemera danica : a new method for estimating degree-days required for growth. Journal of Animal Ecology, 54, 919-930.

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